COSEWIC Assessment and Status Report on the Red Crossbill (Percna) in Canada
- Assessment Summary
- Executive Summary
- COSEWIC History, Mandate, Membership and Definitions
- Lists of Figures, Tables and Appendices
- Species Information
- Population Sizes and Trends
- Limiting Factors and Threats
- Special Significance of the Species
- Existing Protection or Other Status
- Summary of Status Report
- Technical Summary
- Acknowledgements and Literature Cited
- Biographical Summary of the Report Writers, Authorities Contacted, and Collections Examined
We are grateful for the help, discussions and input of many people in the preparation of this report. They include Alex Mosseler, Dave Fifield, Greg Stroud, Chris Cohlmeyer, Ron Summers, Richard Elliot, Keith Lewis, Joe Brazil, Michel Gosselin, Roger Etcheberry, Bradford Gover, Peter Thomas, John Maunder, Kate Bredin and Mark Elderkin. Much of information contained in this report was derived from a manuscript co-authored with Don Steele, Ian Thompson, Rick West and Alex Mosseler, and we are grateful to them for their input. The Canadian Forest Service first initiated interest in the status of Red Crossbill percna.
Funding for the preparation of this status report was provided by the Canadian Wildlife Service, Environment Canada.
Adkisson, C.S. 1996. Red Crossbill. In A. Poole and F. Gill (Editors). The Birds of North America, No. 256. American Ornithologists' Union, Washington, D.C.
American Ornithologists' Union. 1931. Checklist of North American Birds. 4th Edition. American Ornithologists' Union, Lancaster, PA.
American Ornithologists' Union. 1957. Checklist of North American Birds. 5th Edition. American Ornithologists' Union, Baltimore, MD.
American Ornithologists' Union. 1983. Checklist of North American Birds. 6th Edition. Allen Press, Inc., Lawrence, Kansas.
Austin, O.L. (Editor). 1968. Life Histories of North American Grosbeaks, Buntings, Towhees, Finches, Sparrows and Allies. United States National Museum Bulletin 237. Washington, D.C.
Avery, M. and R. Leslie. 1990. Birds and Forestry. Poyser, London.
Benkman, C.W. 1987a. Crossbill foraging behavior, bill structure, and patterns of food profitability. Wilson Bulletin 99: 351 - 368.
Benkman, C.W. 1987b. Food profitability and the foraging ecology of crossbills. Ecological Monographs 57: 251 - 267.
Benkman, C.W. 1988. On the advantages of crossed mandibles: an experimental approach. Ibis 130: 288 - 293.
Benkman, C.W. 1989. On the evolution and ecology of island populations of crossbills. Evolution 43: 1324 - 1330.
Benkman, C.W. 1990. Intake rates and the timing of crossbill reproduction. Auk 17: 376 - 386.
Benkman, C.W. 1992a. A crossbill's twist of fate. Natural History 101: 38 - 42.
Benkman, C.W. 1992b. White-winged Crossbill. In A. Poole and F. Gill (eds.). The Birds of North America, No. 27. American Ornithologists' Union, Washington, D.C.
Benkman, C. W. 1993a. Adaptation to single resources and the evolution of crossbill (Loxia) diversity. Ecological Monographs 63: 305 - 325.
Benkman, C.W. 1993b. Logging, conifers, and the conservation of crossbills. Conservation Biology 7: 473 - 479.
Benkman, C.W. 1993c. The evolution, ecology, and decline of the Red Crossbill of Newfoundland. American Birds 47: 225 - 229.
Benkman, C.W. 1994. Comments on the ecology and status of the Hispaniolan Crossbill (Loxia leucoptera megaplaga), with recommendations for its conservation. Caribbean Journal of Science 30: 250 - 254.
Benkman, C.W. and A.K. Lindholm. 1991. The advantages and evolution of a morphological novelty. Nature 349: 519 - 520.
Benkman, C.W., W.C. Holimon, and J.W. Smith. 2001. The influence of a competitor on the geographic mosaic of coevolution between crossbills and Lodgepole Pine. Evolution 55: 282 - 294.
Bent, A.C. 1912. A new subspecies of crossbill from Newfoundland. Smithsonian Miscellaneous Collections 60: 1 - 3.
Bock, C.E. and L.W. Lepthien. 1976. Synchronous eruptions of boreal seed-eating birds. American Naturalist 110: 559 - 571.
Bock, C.E. and T.L. Root. 1981. The Christmas Bird Count and avian ecology. Studies in Avian Biology 6: 17 - 23.
Brewer, D., A.W. Diamond, E.J. Woodsworth, B.T. Collins, and E.H. Dunn. 2000. Canadian Atlas of Bird Banding. Volume 1: Doves, Cuckoos, and Hummingbirds through Passerines. Special Publication. Canadian Wildlife Service, Ottawa.
Carroll, A.L. 1996. Evaluation of the predictive capacity of the Eastern Hemlock Looper Decision Support System. Pp. 9 - 14 in T.L. Shore and D.A. MacLean (eds.). Proceedings: Decision Support Systems for Forest Pest Management. Forestry Research Development and Applications Report 260.
CITES. 2003. Convention on International Trade in Endangered Species of Wild Flora and Fauna Species Database [Online]. http://www.cites.org/index.html. [Accessed 28/2/2003].
Damman, A.W.H. 1983. An ecological subdivision of the island of Newfoundland. Pp. 163 - 206 in G.R. South (Editor). Biogeography and Ecology of the Island of Newfoundland. Dr W. Junk Publishers, The Hague.
DeBenedictis, P.A. 1995. Red Crossbills, one through eight. Birding 27: 494 - 501.
Dickerman, R.W. 1986. A review of the Red Crossbill in New York State - Part 2. Identification of specimens from New York. Kingbird 36: 127 - 134.
Dickerman, R.W. 1987. The "Old Northeastern" subspecies of Red Crossbill. American Birds 41: 189 - 194.
Dunn, E.H. and J.R. Sauer. 1997. Monitoring Canadian bird populations with winter counts. Pp. 49 - 55 in Monitoring Bird Populations: The Canadian Experience. Canadian Wildlife Service Occasional Paper Number 95.
English, B. 1998. Impact of squirrel predation on White Pine cone collection efforts in Newfoundland. Newfoundland Forest Service Silviculture Notebook 42.
Erskine, A.J. 1977. Birds in boreal Canada: communities, densities and adaptations. Canadian Wildlife Service Report Series. Report 41.
Erskine, A.J. 1992. Atlas of Breeding Birds of the Maritime Provinces. Nimbus Publishing Ltd. and the Nova Scotia Museum, Halifax.
Flight, G.R. and R.D. Peters. 1992. Twenty Year Forestry Development Plan 1990 - 2009. Newfoundland Forest Service, St. John's.
Global Forest Watch Canada. 2000. Canada's Forests at a Crossroads: An Assessment in the Year 2000. World Resources Institute, Washington, D.C.
Godfrey, W.E. 1966. The Birds of Canada. Biological Series No. 73. National Museum of Canada, Ottawa.
Godfrey, W.E. 1986. The Birds of Canada. Revised Edition. National Museum of Natural Sciences (Canada), Ottawa.
Goudie, R.I. 1978. Red Squirrels, Tamiasciurus hudsonicus, in the Salmonier River Valley, Newfoundland. Canadian Field Naturalist 92: 193 - 194.
Griscom, L. 1937. A monographic study of the Red Crossbill. Proceedings of the Boston Society of Natural History 41: 77 - 210.
Griscom, L. and D.E. Snyder. 1955. The Birds of Massachusetts. Peabody Museum, Salem, Massachusetts.
Groth, J.G. 1988. Resolution of cryptic species in Appalachian Red Crossbills. Condor 90: 745 - 760.
Groth, J.G. 1993a. Call matching and positive assortative mating in Red Crossbills. Auk 110: 398 - 401.
Groth, J.G. 1993b. Evolutionary differentiation in morphology, vocalizations, and allozymes among nomadic sibling species in the North American Red Crossbill (Loxia curvirostra) complex. University of California Publications in Zoology 127: 1 - 143.
Hahn, T.P. 1995. Integration of photoperiodic and food cues to time changes in reproductive physiology by an opportunistic breeder, the Red Crossbill, Loxia curvirostra (Aves: Carduelinae). Journal of Experimental Zoology 272: 213 - 226.
Hayward, F.R., J.D. Roberts, W.J. Johnston, P.S. Murray, A.R. Penney, and M.C. Vardy. 1960. Golden Jubilee Report. Newfoundland Forest Protection Association, St. John's.
Hayward, F.R., J.D. Roberts, W.J. Johnston, S.S. Peters, and M.C. Vardy. 1961. Annual Report. Newfoundland Forest Protection Association, St. John's.
Helle, P. 1985. Effects of forest fragmentation on bird densities in northern boreal forests. Ornis Fennica 62: 35 - 41.
Holimon, W.C., C.W. Benkman, and M.F. Willson. 1998. The importance of mature conifers to red crossbills in southeast Alaska. Forest Ecology and Management 102: 167 - 172.
IUCN. 2002. 2002 IUCN Red List of Threatened Species [Online]. http://www.redlist.org/. [Accessed 28/2/2003].
James, P.C., T.W. Barry, A.R. Smith, and S.J. Barry. 1987. Bill crossover ratios in Canadian crossbills Loxia spp. Ornis Scandinavica 18: 310 - 312.
Kaufman, K. 1999. Birds. Audubon 101: 89 - 91
Knox, A.G. 1992. Species and pseudospecies: the structure of crossbill populations. Biological Journal of the Linnean Society 47: 325 - 335.
Koenig, W.D. and J.M.H. Knops. 2001. Seed-crop size and eruptions of North America boreal seed-eating birds. Journal of Animal Ecology 70: 609 - 620.
Lack, D. 1944. Correlation between beak and food in the crossbill, Loxia curvirostra Linnaeus. Ibis 86: 552 - 553.
Lamberton, R.D. 1976. Avifaunal Survey of Gros Morne National Park. Parks Canada Contract Report. Report 74-46.
Lewis, K.P. 1997. Summary of Red Pine Surveys from January 18 to April 12, 1997. Biopsychology Program, Memorial University of Newfoundland, St. John's. Unpublished report.
Lewis, K.P. 1999. Nest Predation in Riparian Buffer Strips in a Balsam Fir Forest in Western Newfoundland. M.Sc. Thesis, Biospsychology Program, Memorial University of Newfoundland, St. John's.
Létourneau, V. 1996. Red Crossbill. Pp. 1066 - 1069 in J. Gauthier and Y. Aubry (Editors). The Breeding Birds of Québec: Atlas of the Breeding Birds of Southern Québec. Association Québécoise des Groupes D'Ornithologues, Province of Quebec Society for the Protection of Birds, Canadian Wildlife Service, Environment Canada, Québec Region, Montreal.
Link, W.A., and J.R. Sauer. 1998. Estimating population change from count data: application to the North American Breeding bird Survey. Ecological Applications 8: 258-268.
Mactavish, B., J.E. Maunder, and W.A. Montevecchi. 1989. Field checklist of the birds of insular Newfoundland and its continental shelf waters. Osprey 20: 139 - 144.
Mactavish, B., J.E. Maunder, and W.A. Montevecchi. 1999. Field checklist of the birds of insular Newfoundland and its continental shelf waters. Osprey 30: 22 -27.
Mactavish, B., J.E. Maunder, W.A. Montevecchi, J.L. Wells and D. Fifield. 2003. Checklist of the birds of insular Newfoundland and its continental shelf waters. Natural History Society of Newfoundland and Labrador, St. John’s.
Mannion, J.J. (Editor). 1977. The Peopling of Newfoundland. Institute for Social and Economic Research Memorial University of Newfoundland Paper Number 8.
Marquiss, M. and R. Rae. 1994. Seasonal trends in abundance, diet and breeding of Common Crossbills (Loxia curvirostra) in an area of mixed species conifer plantation following the 1990 crossbill 'irruption'. Forestry 67: 31 - 47.
Marquiss, M. and R. Rae. 2002. Ecological differentiation in relation to bill size amongst sympatric, genetically undifferentiated crossbills Loxia spp. Ibis 144: 494‑508.
Maunder, J.E. and W.A. Montevecchi. 1982. Field checklist of the birds of insular Newfoundland. Osprey 13: 59 - 62.
McCabe, T.T. and E.B. McCabe. 1933. Notes on the anatomy and breeding habits of crossbills. Condor 35: 136 - 147.
McLaren, I.A. 1991. Northeastern regional report. American Birds 45: 244 - 247.
Minty, D. 1976. Red Squirrels in Newfoundland. Osprey 7: 19 - 24.
Montevecchi, W.A. and L.M. Tuck. 1987. Newfoundland Birds: Exploitation, Study, Conservation. Publications of the Nuttall Ornithological Club, No. 21. Nuttall Ornithological Club, Cambridge, Massachusetts.
Montevecchi, W.A. and J. Wells. 1987. Vernacular bird names of Newfoundland. Pp. 232 - 245 in W. A. Montevecchi and L.M. Tuck. Newfoundland Birds: Exploitation, Study, Conservation. Nuttall Ornithological Club, Cambridge, MA.
National Audubon Society. 2002. The Christmas Bird Count Historical Results [Online]. http://www.audubon.org/bird/cbc. [Accessed 31/1/2003].
Natural Resources Canada. 2000. Geogratis [Online]. http://geogratis.gc.ca/. [Accessed 30/1/2003].
Natural Resources Canada. 2002. The State of Canada's Forests: Reflections of a Decade 2001 - 2002 [Online]. Government of Canada. http://www.nrcan.gc.ca/cfs-scf/national/what-quoi/sof/latest_e.html. [Accessed 24/3/2003].
NatureServe Explorer: An online encyclopedia of life. 2002. Version 1.6. NatureServe, Arlington, Virginia. http://www.natureserve.org/explorer. [Accessed 28/2/2003].
Newton, I. 1972. Finches. Collins, London.
Nf.birds Newsgroup Archives. 2003. [Accessed 31/1/2003].
Noble, G.K. 1919. Notes on the avifauna of Newfoundland. Bulletin of the Museum of Comparative Zoology 62: 543 - 568.
Ouellet, H. 1969. Les oiseaux de l'île Anticosti, province de Québec, Canada. National Museum of Natural Sciences Publications in Zoology, No. 1. National Museums of Canada.
Page, G., W. C. Wilton, and T. Thomas. 1974. Forestry in Newfoundland. Newfoundland Forest Research Centre, St. John's, Newfoundland.
Parchman, T.L. and C.W. Benkman. 2002. Diversifying coevolution between crossbills and Black Spruce on Newfoundland. Evolution 56: 1663 - 1672.
Payne, N.F. 1976. Red Squirrel introduction to Newfoundland. Canadian Field Naturalist 90: 60 - 64.
Payne, R.B. 1987. Populations and type specimens of a nomadic bird: comments on the North American crossbills Loxia pusilla Gloger 1834 and Crucirostra minor Brehm 1845. Occasional Papers of the Museum of Zoology University of Michigan 714: 1 - 37.
Peters, H.S. and T.D. Burleigh. 1951. The Birds of Newfoundland. Newfoundland Department of Natural Resources, St. John’s.
Pimm, S.L. 1990. The decline of the Newfoundland crossbill. Trends in Ecology and Evolution 5: 350 - 351.
Pitocchelli, F.J. 1981. Tape Recording of Newfoundland Avian Vocalizations. Centre for Newfoundland Studies, Memorial University of Newfoundland, St. John's.
Pyle, P. 1997. Identification Guide to North American Birds. Volume 1. Slate Creek Press, Bolinas, CA.
Rajora, O.P., L. DeVerno, A. Mosseler, and D.J. Innes. 1998. Genetic diversity and population structure of disjunct Newfoundland and central Ontario populations of Eastern White Pine (Pinus strobus). Canadian Journal of Botany 76: 500 - 508.
Reeks, H. 1869. Notes on the zoology of Newfoundland, letters 1 - 4, ornithology. Zoologist 4: 1609 - 1614, 1689 - 1695, 1741 - 1759, 1849 - 1858.
Robb, M.S. 2000. Introduction to vocalizations of crossbills in north-western Europe. Dutch Birding 22: 61 - 107.
Roberts, B.A. 1985. Distribution and extent of Pinus resinosa Ait. in Newfoundland. Rhodora 87: 341 - 356.
Roberts, B.A. and A.U. Mallik. 1994. Responses of Pinus resinosa in Newfoundland to wildfire. Journal of Vegetation Science 5: 185 - 196.
Rooke, K.B. 1935. Observations on the birds of Newfoundland during the 1934 expedition of the Public Schools Exploring Society. Ibis 5: 856 - 879.
Ryan, A.G. 2003. Protected Areas in Newfoundland & Labrador [Online]. Natural Areas, Department of Forest Resources & Agrifoods, Government of Newfoundland & Labrador. http://www.gov.nf.ca/forestry/protection/areas.asp. [Accessed 28/2/2003].
Sauer, J.R., Hines, J.E., and Fallon, J. 2001. The North American Breeding Bird Survey, Results and Analysis 1966 - 2000 [Online]. Version 2001.2. USGS Patuxent Wildlife Research Center, Laurel, MD. http://www.mbr-pwrc.usgs.gov/bbs/. [Accessed 28/2/2003].
Sibley, D.A. 2000. The Sibley Guide to Birds. Alfred A. Knopf Inc., New York.
Smith, J.W., C.W. Benkman, and K. Coffey. 1999. The use and misuse of public information by foraging Red Crossbills. Behavioral Ecology 10: 54 - 62.
Smith, P.W. 1997. The history and taxonomic status of the Hispaniolan Crossbill Loxia megaplaga. Bulletin of the British Ornithologists' Club 117: 264 - 271.
Snyder, D.E. 1954. A nesting study of Red Crossbills. Wilson Bulletin 66: 32 - 37.
Summers, R.W. 2002. Cone sizes of Scots pines Pinus sylvestris in the Highlands of Scotland implications for pine-eating crossbills Loxia spp. in winter. Forest Ecology and Management 164: 303 - 305.
Summers, R.W., D.C. Jardine, M. Marquiss, and R. Rae. 2002. The distribution and habitats of crossbills Loxia spp. in Britain, with special reference to the Scottish Crossbill Loxia scotica. Ibis 144: 393 - 410.
Summers, R.W., and R. Proctor. 1999. Tree and cone selection by crossbills Loxia sp. and red squirrels Sciurus vulgaris at Abernethy forest, Strathspey. Forest Ecology and Management 118: 173 - 182.
Summers, R.W., and S.B. Piertney. 2003. The Scottish Crossbill – what we know and what we don’t. British Birds 96: 100 - 111.
Thompson, I.D., H.A. Hogan, and W.A. Montevecchi. 1999. Avian communities of mature Balsam Fir forests in Newfoundland: age-dependence and implications for timber harvesting. Condor 101: 311 - 323.
Thompson, I.D., D.J. Larson, and W.A. Montevecchi. 2003. Characterization of old "wet boreal" forests, with an example from balsam fir forests of western Newfoundland. Environmental Reviews 11:S23 - S46.
Todd, W.E.C. 1963. Birds of the Labrador Peninsula and Adjacent Areas. Carnegie Museum and the University of Toronto Press, Pittsburgh, PA.
Tordoff, H.B. and W.R. Dawson. 1965. The influence of day length on reproductive timing in the Red Crossbill. Condor 67: 416 - 422.
Tuck, L.M. and M.J. Borotra. 1972. Additions to the Avifauna of St. Pierre and Miquelon. Canadian Field Naturalist 86: 279 - 284.
Tufts, R.W. 1986. The Birds of Nova Scotia. 3rd Edition. Nimbus Publishing and the Nova Scotia Museum, Halifax, Nova Scotia.
US Fish & Wildlife Service. 2003. The Endangered Species Program [Online]. http://endangered.fws.gov/. [Accessed 28/2/2003].
USGS Patuxent Wildlife Research Center. 2003. North American Breeding Bird Survey Internet data set [Online]. http://www.mp2-pwrc.usgs.gov/bbs/index.html. [Accessed 24/1/2003].
Wells, J. and W.A. Montevecchi. 1984. The birds of Baccalieu Island. Osprey 15(2): 34 – 40.
West, R.J. 1989. Cone depredations by the Red Squirrel in Black Spruce stands in Newfoundland: implications for commercial cone collection. Canadian Journal of Forest Research 19: 1207 - 1210.
Whitaker, D.M., W.A. Montevecchi, and A. Mosseler. 1996. Red Pine, wildlife and Scleroderris canker in Newfoundland. Osprey 27: 64 - 71.
Wilton, W.C. and C.H. Evans. 1974. Newfoundland Forest Fire History, 1619 - 1960. Forestry Canada Newfoundland and Labrador Information Report. Newfoundland Forest Research Centre, Report N-X-116.
Wren, L.S. 2001. Continental and Regional Distribution and Abundance Patterns of Boreal Cardueline Finches: Influences of Conifer Seed Availability. M.Sc. Thesis, Biopsychology Program, Memorial University of Newfoundland, St. John's, NF.
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